↑Esselstyn, J.A., Garcia, H.J.D., Saulog, M.G. and Heaney, L.R. 2008. A new species of Desmalopex (Pteropodidae) from the Philippines, with a phylogenetic analysis of the Pteropodini. Journal of Mammalogy 89(4):815–825.
↑Giannini, N.P., Almeida, F.C., Simmons, N.B. and Helgen, K.M. 2008. The systematic position of Pteropus leucopterus and its bearing on the monophyly and relationships of Pteropus (Chiroptera: Pteropodidae). Acta Chiropterologica 10(1):11–20.
